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In both types of feather, the barbules that extend from the barb ramus grow from a single layer of cells, called the barbule plate, on the periphery of the barb ridge.

而不论是何种，主延出来的，脊表面叫做「板」的单层细胞长出来。

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原声例句
大英百科全书

Each barb branches into barbules, which link to one another by hooks.

每根羽支分拨成羽，羽通过钩子相互连接。

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Space – Verge Science

And lots of times, there are little feather barbules in there that we can look at the microscopic characters

很多时候， 里有羽，我看到微观特征。

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PBS地球-动物趣闻

The central part of the feather became a hardened structure called a rachis, with lots of little barbs and barbules, creating the vanes of the feather.

羽的中央部分变成了一种称为羽轴的硬化结构，上有许多倒钩和枝，形成了羽的叶片。

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