The carpel is homologous with the megasporophylls of certain spore-bearing vascular plants (pteridophytes) and the ovuliferous scales of gymnosperms.

心皮和能产生孢子维管植（羊齿类）还有珠子植孢子叶是同源。

Bract scale The structure found in large numbers in the female cone of gymnosperms, each bearing an ovuliferous scale in its axil.

在子植雌球果中量存在结构，每个苞鳞都包含一个珠鳞。